The Graduate School of Arts and SciencesBiomedical Programs2020 – 2021The Graduate Bulletinfor Graduate Students in Biomedical Graduate ProgramsAdministered on the School of Medicine Campus

On the Cover: PhD student Andy Kwok (Integrative Physiology and Pharmacology)displays patience in a research laboratory.

Table of Contents2020-2021 Academic Calendar . 6Programs of Study . 7The University . 10The Administration. 18The Graduate School . 21Procedures . 27Admissions . 27How to Apply . 27Eligibility . 27Students with Disabilities . 27Admission Categories . 28Classification of Admitted Students . 28Cost of Attendance . 29Tuition Schedule . 29Fee Schedule . 29Tuition Concession . 29Tuition for Courses Taken in Reynolda campus-based graduate programs . 29Student Graduate Fees . 30Past Due Balances . 30Tuition Refunds Due to Withdrawal . 30Financial Aid. 31Satisfactory Academic Progress . 31Graduation At-Risk Status. 31Academic Probation. 31Appeal to Financial Aid Ineligibility Status . 32Student Loans . 32Graduate School Scholarship Awards . 32Academic & Coursework Practices .33Statement on Student Rights and Responsibilities. 33Honor Code & Non-Academic Code of Conduct . 33Grievance Procedures . 33Academic Grievance Procedures . 34Registration Practices . 37Term Registration . 37Course Registration . 37Grading Practices . 39Changes in Status. 40Student Wellness . 43Health Service . 43Counseling Services . 45

Requirements for Degrees . 46Degrees Offered . 46Thesis/Dissertation Committee Composition & Review Guidelines . 46Final Examination Assessment . 47Requirements for the Master of Science . 47Requirements for the Master of Healthcare Leadership . 49Requirements for the Doctor of Philosophy . 50Degree Programs. 52Graduate School General Studies (GRAD). 52Other Graduate Coursework Available Through Reynolda-campus Based Programs of Study . 56Addiction Research and Clinical Health (ARCH) .59Biochemistry and Molecular Biology (BAMB) . 63Biomedical Engineering (BMES) . 66Biomedical Science (BMSC) . 73Cancer Biology (CABI) . 78Clinical Research Management (CRM) . 82Comparative Medicine (COMD) . 88Genetic Counseling (GENC) . 90Health Disparities in Neuroscience-related Disorders (HDND) . 94Healthcare Leadership (HCL) .95Integrative Physiology and Pharmacology (IPP) .100Microbiology and Immunology (MICR) . 105Molecular and Cellular Biosciences (MCB) . 108Molecular Genetics and Genomics (MOGN) . 113Molecular Medicine and Translational Science (MMTS) . 116Neuroscience (NEUR) . 122Translational and Health System Science (THSS) . 132Joint Degrees, Certificates, and Concentrations. 140Joint Degrees . 140PhD/MMS (Molecular Medicine and Translational Science) . 140PhD/MD . 141PhD/MBA .143MS/MD (Translational & Health System Science) . 145MS/BS-BA (Neuroscience) . 145Graduate Certificates . 147Clinical and Translational Investigation . 147

Learning Health System Science . 147Concentrations . 149Antimicrobial Stewardship and Infection Prevention.149Cancer Biology .150Integrative Physiology and Pharmacology. 151Neuroscience . 151Regenerative Medicine . 152

2020-2021 Academic CalendarSUMMER TERM 2020MayMayJuneJuneJulyAugustAugustAugust20213116 – 1771414AugustAugustSeptember15-20284FALL TERM 2020August20August21August26September 7September 15September 16November 9 – 20November 25 – 27December 15December 17December 18-23December 22JanuaryJanuary815SPRING TERM ruary2March8 – 12March 22 – April 1April2April20April28April 29 – May 4May5MayMayMay101517New Student OrientationClasses BeginDeadline: Statement of Intent form to Grad School for Sept graduatesLast day to add/drop a courseCourse Registration for Fall 2020Deadline: Last day to defend for September graduatesClasses EndDeadline: ETD Student Advisor Agreement, final copy of thesis/dissertation, exit survey completions for September graduatesExaminationsGrades DueGraduation (Degrees Conferred)New Student OrientationProgram Orientation/Advising MeetingsClasses BeginLABOR DAY HOLIDAYDeadline: Statement of Intent form to Grad School for Jan graduatesLast day to add or drop a courseCourse Registration for Spring 2021THANKSGIVING HOLIDAYDeadline: Last day to defend for January graduatesClasses EndExaminationsDeadline: ETD Student Advisor Agreement, final copy of thesis/dissertation, exit survey completions for January graduatesGrades DueGraduation (Degrees Conferred)New Student OrientationProgram Orientation/Advising MeetingsClasses beginMARTIN LUTHER KING JR. HOLIDAYDeadline: Statement of Intent form to Grad School for May graduatesLast day to add or drop a courseSPRING BREAKCourse Registration for Summer 2021GOOD FRIDAY HOLIDAYDeadline: Last day to defend for May graduatesClasses endExaminationsDeadline: ETD Student Advisor Agreement, final copy of thesis/dissertation, exit survey completions for May graduatesGrades DueHooding and Awards CeremonyCommencement

Programs of StudyThe Graduate School administers programs on the Wake Forest School of Medicinecampus and the Reynolda campus, home of the undergraduate college and theirassociated graduate programs.On the School of Medicine campus, the Graduate School offers 8 PhD programs inthe biomedical sciences, MS programs in 11 disciplines, and graduate certificates in 3disciplines. All programs of study, joint degree programs, certificates, andconcentrations subject to this Graduate Bulletin are noted in bold below. Allother programs, including the PhD programs in biology, chemistry, and physics, andMaster’s programs in 22 disciplines, are administered by the Graduate School officebased on the Reynolda campus and covered by a separate Graduate Bulletin. TheGraduate School also offers 15 joint degree programs in conjunction with the WakeForest School of Medicine, School of Business, School of Divinity, and School of Law; aswell as 13 certificate programs. Contact information on all programs and certificates ofstudy may be found on our website at Programs of StudyBiology (PhD)Biomedical Engineering (PhD)Chemistry (PhD)Integrative Physiology and Pharmacology (PhD)Molecular & Cellular Biosciences(a non-degree granting admissions pathway to the following degree granting programs) Biochemistry and Molecular Biology (PhD) Cancer Biology (PhD) Microbiology and Immunology (PhD) Molecular Genetics and Genomics (PhD) Molecular Medicine and Translational Science (PhD)Neuroscience (PhD)Physics (PhD)Master’s Programs of StudyAddiction Research and Clinical Health (MS)Bioethics (MA)Biology (MS)Biomedical Engineering (MS)Biomedical Science (MS)Chemistry (MS)Clinical Research Management (MS - online)Communication (MA)

Comparative Medicine (MS)Computer Science (MS)Counseling (MA)Counseling (MA - online)Counseling - Human Services (MAHS - online)Documentary Film (MA/MFA)Education (MAEd)English (MA)Genetic Counseling (MS)Health and Exercise Science (MS)Healthcare Leadership (MHL - online)Health Disparities in Neuroscience-related Disorders (MS)Interpreting and Translation Studies (MA): Interpreting and Translation Studies Intercultural Services in Healthcare Teaching of InterpretingLiberal Arts Studies (MA)Mathematics & Statistics (MA)Molecular Medicine and Translational Science (MS)Neuroscience (MS)Physics (MS)Psychology (MA)Religious Studies (MA)Sustainability (MA)Translational & Health System Science (MS)Joint Degree ProgramsMS & BS/A (Bioethics)MS & BS/A (Computer Science)MS & BS/A (Neuroscience)MA/JD (Bioethics)MA/JD (Religious Studies)MA/JD (Sustainability)MA/MD (Bioethics)MS/MD (Translational and Health System Science)PhD/MDMA/MDiv (Bioethics)MA/MDiv (Counseling)MAEd/MDiv (Education)MA/MDiv (Sustainability)PhD/MMS (MMTS & PA)PhD/MBA

Graduate CertificatesBioethics: Bioethics Biomedical Research Ethics Clinical BioethicsClinical and Translational InvestigationCurriculum, Instruction, and AssessmentInterpreting and Translation Studies: Intercultural Services in Healthcare Interpreting Studies Teaching of Interpreting (Postgraduate) Translation StudiesLearning Health System ScienceMedieval and Early Modern StudiesStructural and Computational l Stewardship & Infection PreventionCancer BiologyIntegrative Physiology and PharmacologyNeuroscienceRegenerative MedicineReligion and Public EngagementWomen’s Gender, and Sexuality Studies

The UniversityWake Forest University is characterized by its devotion to liberal learning andprofessional preparation for men and women, its strong sense of community andfellowship, and its encouragement of free inquiry and expression.Wake Forest Institute was founded in 1834 by the Baptist State Convention of NorthCarolina. The school opened its doors on February 3 with Samuel Wait as principal.Classes were first held in a farmhouse on the Calvin Jones plantation in Wake County,North Carolina, near which the village of Wake Forest later developed.Rechartered in 1838 as Wake Forest College, Wake Forest is one of the oldestinstitutions of higher learning in the state. The School of Law was established in 1894,followed by a two-year medical school in 1902. Wake Forest was exclusively a college formen until World War II, when women were admitted for the first time.In 1941 the medical school moved to Winston-Salem to become affiliated with NorthCarolina Baptist Hospital and was renamed the Bowman Gray School of Medicine. In1946 the trustees of Wake Forest and the Baptist State Convention of North Carolinaaccepted a proposal by the Z. Smith Reynolds Foundation to relocate the College toWinston-Salem.The late Charles and Mary Reynolds Babcock donated much of the R.J. Reynoldsfamily estate as the site for the campus and building funds were received from manysources. From 1952 to 1956, the first fourteen buildings were constructed in Georgianstyle on the new campus. The move to Winston-Salem took place in the summer of1956; the original, or “old” campus, is now home to Southeastern Baptist TheologicalSeminary.Following the move, Wake Forest grew considerably in enrollment, programs, andstature and became a University in 1967. The School of Business Administration, firstestablished in 1948, was named the Charles H. Babcock School of BusinessAdministration in 1969 and admitted its first graduate students in 1971. In 1972 theschool enrolled only graduate students and the name was changed to the Charles H.Babcock Graduate School of Management; departments of business and accountancyand economics were established in the College. In 1980 the Department of Business andAccountancy was reconstituted as the School of Business and Accountancy; the namewas changed to the Wayne Calloway School of Business and Accountancy in 1995. OnJuly 1, 2009, the Wayne Calloway School of Business and Accountancy and the CharlesH. Babcock Graduate School of Management officially merged under the name WakeForest University Schools of Business.The Division of Graduate Studies, established in 1961, is now organized as theGraduate School and encompasses advanced work in the arts and sciences on both theReynolda and Bowman Gray campuses. In 1997 the medical school was renamed theWake Forest School of Medicine; its campus is now known as the Bowman GrayCampus. The School of Divinity was established in 1999.Wake Forest honors its Baptist heritage in word and deed. The University will fulfillthe opportunities for service arising out of that heritage. Governance is by an

independent Board of Trustees; there are advisory boards of visitors for the College andeach professional school. A joint board of University trustees and trustees of the NorthCarolina Baptist Hospital is responsible for Wake Forest University Baptist MedicalCenter, which includes the hospital and the medical school.The College, School of Business, School of Law, and the School of Divinity arelocated on the Reynolda Campus in northwest Winston-Salem and the MorrocroftCampus in Charlotte, NC. The Wake Forest School of Medicine is about five miles away,with locations in the city’s downtown area and in the Baptist Medical Center. TheGraduate School maintains operations on both the Reynolda and School of Medicinecampuses depending on the program of study. The University also offers instructionregularly at Casa Artom in Venice, at Worrell House in London, at Flow House inVienna, and in other places around the world.The College offers courses in more than forty fields of study leading to thebaccalaureate degree. The School of Divinity offers the Master of Divinity degree. TheWake Forest School of Business offers a four-year bachelor of science degree, withmajors in accountancy, business and enterprise management, finance, andmathematical business (offered jointly with the Department of Mathematics); and threegraduate degree programs, Master of Science in Accountancy (MSA), Master of Arts inManagement (MA), and Master of Business Administration (MBA).The School of Law offers the Juris Doctor and Master of Laws in American lawdegrees. The school also offers a joint JD/MBA degree with the Schools of Business. TheWake Forest School of Medicine offers the Doctor of Medicine degree (MD) as well asthe Master of Medical Science (MMS) through the Physician Assistant program, and theMaster of Science (MS) in Nurse Anesthesia, and the Doctor of Nursing Practice (DNPdegree). The School of Medicine and the Schools of Business offer a joint MD/MBAprogram.The Graduate School confers the Master of Arts, Master of Arts in Education, Masterof Arts in Human Services, Master of Arts in Liberal Studies, Master of Fine Arts, andMaster of Science degrees in the arts and sciences and biomedical science, and theDoctor of Philosophy degree. The Graduate School also offers an MD/MS, MD/PhD,PhD/MMS, as well as an MD/MA in bioethics jointly with the School of Medicine, and aPhD/MBA program jointly with the School of Business. In addition, the GraduateSchool offers an MDiv/MA in bioethics, counseling, or sustainability as well as aMDiv/MAED jointly with the School of Divinity; and a JD/MA in bioethics, religiousstudies or sustainability jointly with the School of Law. Certificates are offered inBioethics, Clinical and Translational Investigation, Learning Health System Science,Interpreting and Translation Studies, Structural and Computational Biophysics, andSustainability.STATEMENT OF PURPOSE OF THE UNIVERSITYFollowing is the official statement of the purposes and objectives of the University:Wake Forest is a University dedicated to the pursuit of excellence in the liberal artsand in graduate and professional education. Its distinctiveness in its pursuit of itsmission derives from its private, coeducational, and residential character; its size and

location; and its Baptist affiliation. Each of these factors constitutes a significantaspect of the unique character of the institution. The University is comprised of WakeForest College, the Graduate School, the School of Law, the School of Medicine, theSchool of Divinity and the School of Business. It seeks to honor the ideals of liberallearning, which entail commitment to transmission of cultural heritages; teaching themodes of learning in the basic disciplines of human knowledge; developing the criticalappreciation of moral, aesthetic, and religious values; advancing the frontiers ofknowledge through in-depth study and research; and applying and using knowledge inthe service of humanity.Wake Forest has been dedicated to the liberal arts for over a century and a half; thismeans education in the fundamental fields of human knowledge and achievement, asdistinguished from education that is technical or narrowly vocational. It seeks toencourage habits of mind that ask “why,” that evaluate evidence, that are open to newideas, that attempt to understand and appreciate the perspectives of others, that acceptcomplexity and grapple with it, that admit error, and that pursue truth. Wake ForestCollege has by far the largest student body in the University, and its function is centralto the University’s larger life. The College and the Graduate School are most singularlyfocused on learning for its own sake; they, therefore, serve as exemplars of specificacademic values in the life of the University.Beginning as early as 1894, Wake Forest accepted an obligation to provideprofessional training in a number of fields as a complement to its primary mission ofliberal arts education. This responsibility is fulfilled in the conviction that the humanevalues embodied in the liberal arts are also centrally relevant to the professions.Professional education at Wake Forest is characterized by a commitment to ethical andother professional ideals that transcend technical skills. Like the Graduate School, theprofessional schools are dedicated to the advancement of learning in their fields. Inaddition, they are specifically committed to the application of knowledge to solvingconcrete problems of human beings. They are strengthened by values and goals whichthey share with the College and Graduate School, and the professional schools enhancethe work of these schools and the University as a whole by serving as models of serviceto humanity.Wake Forest was founded by private initiative, and ultimate decision-makingauthority lies in a privately appointed Board of Trustees rather than in a public body.Funded to a large extent from private sources of support, Wake Forest is determined tochart its own course in the pursuit of its goals. As a coeducational institution it seeks to“educate together” persons of both sexes and from a wide range of backgrounds—racial,ethnic, religious, geographical, socioeconomic, and cultural. Its residential features areconducive to learning and to the pursuit of a wide range of co-curricular activities. It hasmade a conscious choice to remain small in overall size; it takes pride in being able tofunction as a community rather than a conglomerate. Its location in the Triad area ofNorth Carolina engenders an ethos that is distinctively Southern, and more specificallyNorth Carolinian. As it seeks to broaden further its constituency and to receive nationalrecognition, it is also finding ways to maintain the ethos associated with its regionalroots.

Wake Forest is proud of its Baptist and Christian heritage. For more than a centuryand a half, it has provided the University an indispensable basis for its mission andpurpose, enabling Wake Forest to educate thousands of ministers and lay people forenlightened leadership in their churches and communities. Far from being exclusive andparochial, this religious tradition gives the University roots that ensure its lastingidentity and branches that provide a supportive environment for a wide variety of faiths.The Baptist insistence on both the separation of church and state and local autonomyhas helped to protect the University from interference and domination by outsideinterests, whether these be commercial, governmental, or ecclesiastical. The Baptistemphasis upon revealed truth enables a strong religious critique of human reason, evenas the claims of revelation are put under the scrutiny of reason. The character ofintellectual life at Wake Forest encourages open and frank dialogue and providesassurance that the University will be ecumenical and not provincial in scope, and that itmust encompass perspectives other than the Christian. Wake Forest thus seeks tomaintain and invigorate what is noblest in its religious heritage.LIBRARIESThe libraries of Wake Forest University support instruction and research at theundergraduate level and in the disciplines awarding graduate degrees. The libraries ofthe University hold membership in the American Library Association and in theAssociation of Southeastern Research Libraries.The Wake Forest University libraries include the Z. Smith Reynolds (ZSR) Library,which is located on the Reynolda Campus and primarily supports the undergraduateCollege, the Wake Forest School of Business, the Reynolda-based programs of theGraduate School of Arts and Sciences, and the School of Divinity. The ProfessionalCenter Library, housed in the Worrell Professional Center on the Reynolda Campus,serves the School of Law. The Coy C. Carpenter Library primarily serves the WakeForest School of Medicine and the biomedical science programs of the Graduate Schoolof Arts and Sciences and is located on the Bowman Gray Campus.The three library collections total over 2.4 million titles, including over 1.4 million ebooks, more than 100,000 electronic journals and over 15,000 DVDs as well asstreaming media and other formats. The ZSR Library serves as a congressionallydesignated selective federal depository. The Professional Center Library holds nearly130,000 volumes and the Coy C. Carpenter Library holds nearly 27,000 volumes. Thethree libraries share an online search portal, which also provides access to books,electronic resources, journals and databases. Through interlibrary loan service,students, faculty and staff may obtain materials from other libraries at no charge.BOWMAN GRAY CAMPUS AND INNOVATION QUARTER LIBRARY RESOURCESAll faculty, staff and students in the Wake Forest University Graduate School of Artsand Sciences and Biomedical Sciences have full and unrestricted access to the Coy C.Carpenter Library of Wake Forest School of Medicine at its main facility on the firstfloor of the Gray Building. The Library is centrally located within the Wake Forestaffiliated hospital (Wake Forest Baptist Medical Center), and most of the resources are

available online. The main facility of the Library is also well-stocked with print volumescovering all medical and surgical specialties, as well as the basic sciences. There is abranch of the Library located on the third floor of the Bowman Gray Center for MedicalEducation (BGCME) building in Innovation Quarter that is reserved mostly forstudents. The Carpenter Library’s website,, offers access toPubMed, UpToDate, Journal Citation Reports, Micromedex, PsycINFO, and Web ofScience, as well as over 70 other bibliographic and full text databases.Graduate students receive free document delivery service for materials the Libraryowns and interlibrary loan service for materials from other libraries when the Librarydoes not own or license a particular journal or book. In addition, all graduate studentshave unlimited copying and printing services at both locations.T

the biomedical sciences, MS programs in 11 disciplines, and graduate certificates in 3 disciplines. All programs of study, joint degree programs, certificates, and concentrations subject to this Graduate Bulletin are noted in bold below. All other programs, including the PhD